An insight into the structure, composition and hardness of a biological material: the shell of freshwater mussels.

The shell of the freshwater mussel (Mollusca: Bivalvia) is a composite biological material linked with multifunctional roles in sustaining ecosystem services. Apart from providing mechanical strength and support, the shell is an important site for adherence and growth of multiple types of algae and periphyton. Variations in the shell architecture are observed in the mussels both within a species and among different species.