It has been shown that the complex functioning of the baseplate of the bacteriophage T4 is based on the high level hierarchy in the structure organization and on the interaction of protein components forming the hub and the "channels" of short and long fibers. The presence of structure proteins with stabilizing and destabilizing functions has been revealed. It has been shown that the process of binding of long tail fibers with cell receptors is a cooperative process. In general the baseplate can be considered as a logic module providing the transition from the adsorbtion to the nonreversible reorganization in the bacteriophage functioning.

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