A progressive-ratio reinforcement schedule, in which successive reinforcements required an additional 50 responses, was programmed on one key. A response on a second key reset the progressive-ratio schedule to the first step. Before punishment, all pigeons consistently reset the schedule after reinforcement on the first step, thereby minimizing the number of responses required for reinforcement. Punishment was a brief electric shock contingent upon each response on the reset key. The first effect of punishment was to change the frequency of extra responses on the reset key. Under higher intensities of punishment, the pigeons completed the advanced steps of the progressive-ratio schedule before resetting to the first step. Completions of advanced steps were accompanied by decreases in the overall rate of responding and the rate of reinforcement. When the punishment contingency was removed, the major features of pre-punishment performance were recovered.

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http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1338462PMC
http://dx.doi.org/10.1901/jeab.1968.11-133DOI Listing

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