Despite the current recognition of and as two separate valid species of China, neither species have been revised based on examination of their types and/or topotypical materials, nor have they genetically analyzed. In this study, examination of the holotype of showed that it has a serrated anterior edge of the pectoral spine, a slightly emarginate caudal fin, and longer maxillary barbels extending beyond the base of the pectoral spine, the characters shared with specimens currently identified as . Morphological comparisons and molecular analysis showed that specimens from mainland China, which are characterized by the three mentioned morphological features, represent a single species. According to the nomenclatural rule of priority, is a senior subjective synonym of . Within this concept, is widely distributed in the lower reaches of Yangtze River and coastal rivers in Zhejiang and Fujian Province and closely related to , which is endemic to Japan. The morphological differences and species-level genetic distance between and provide additional support for synonymization of and . The paper also describes ontogenetic color changes and coloration polymorphism in this species. Phylogeny of the group, to which belongs, is also discussed.
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