Intergrative Taxonomic Study of the Complex with a Modern Circumscription of the Section (Frullaniaceae, Marchantiphyta).

(subg. ) sect. has been studied using integrative taxonomy methods and utilizing sampling from almost all areas of distribution of the species previously referred to this section. A phylogenetic analysis based on nuclear ribosomal ITS1-2 and chloroplast L-F sequence data and a morphological study reveal a wide range of morphological variability within specimens that has largely disguised the overall taxonomic diversity. , previously regarded as a widespread species, has been found to represent a group of separate species within different sections of subg. : and in sect. , in sect. , and in . sect. II. Illustrations of the type specimens of , , and , as well as illustrations of the sequenced specimens belonging to two of the discussed species ( and ), are provided. The morphological differences separating the highly similar , , , , and are discussed. A dichotomous key is presented for accepted species. New combinations are provided for two taxa.