It was supposed that the current records of might be a species complex composed of multiple species. In this study, on the basis of wide sampling, we found that the records of from Yunnan, China, and northern Vietnam actually represent a cryptic species based on morphological and molecular evidence. can be distinguished from other members of the species group by its moderate body size (SVL 35.5‒37.3 mm in males and 39.2‒45.7 mm in females); white spines on the temporal region, loreal region, snout, and lips in breeding males but absent in females; overlapping heels; tibiotarsal articulation reaching tip of snout; indistinct longitudinal glandular folds on the skin of the shoulders; presence of supernumerary tubercles below the base of fingers II‒IV, distinct pineal body; presence of vomerine teeth; and absence of vocal sacs. Phylogenetic analysis supports that the new species is sister to and the populations from Jingxi, Guangxi and Lào Cai, Vietnam previously reported as also belong to it. Additionally, our results indicate that more cryptic species may exist within the species group, implying that more studies are needed to achieve a complete understanding of the species diversity of this group.

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http://www.ncbi.nlm.nih.gov/pmc/articles/PMC9848789PMC
http://dx.doi.org/10.3897/zookeys.1112.82551DOI Listing

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