Service d'ophtalmologie, CHU de Clermont-Ferrand, 58, rue Montalembert, 63000 Clermont-Ferrand, France; Genetic Reproduction and Development Laboratory (GReD), Translational Approach to Epithelial Injury and Repair Team, CNRS UMR 6293, INSERM U1103, université Clermont-Auvergne, 28, Place Henri Dunant, 63000 Clermont-Ferrand, France.