Currently, contains two subspecies: subsp. and subsp. . In a study by Varsha and Nampoothiri, high (99.7 %) and (99.6 %) sequence similarities indicated that subsp. and probably have a close taxonomic relationship; low (92.2 %) and (97.8 %) sequence similarities and relatively low DNA-DNA hybridization value (75.8 %) indicated that subsp. and subsp. probably represent two different species. In the present study, the taxonomic relationships between subsp. , subsp. and were re-examined based on sequence analyses of 16S rRNA, , , and genes, average nucleotide identity (ANI), digital DNA-DNA hybridization (dDDH) values, average amino acid identity (AAI), fatty acid methyl ester analysis and phenotypic characterization. subsp. LMG 30663 exhibited 97.3 % ANI, 78.3 % dDDH and 96.4 % AAI values to NBRC 109475, higher than the threshold for species demarcation (95-96, 70 and 95-96 %, respectively), indicating that subsp. LMG 30663 and NBRC 109475 belong to the same species. subsp. LMG 30663 had 91.2 % ANI, 43.3 % dDDH and 92.9-93.0% AAI values with the type strain of subsp. , indicating that they represent two different species. Because has been proposed and validated before subsp. , subsp. is transferred to as subsp. comb. nov. The type strain of subsp. is BSN307 (=DSM 100577=MCC 2824=KCTC 21083=LMG 30663). The type strain of subsp. is 516 (=NBRC 109475=BCRC 80576).

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