Three new chasmosaurines from the Kirtland Formation (~75.0-73.4 Ma), New Mexico, form morphological and stratigraphic intermediates between (~74.7-75 Ma, Fruitland Formation, New Mexico) and (~72-71 Ma, Horseshoe Canyon Formation, Alberta). The new specimens exhibit gradual enclosure of the parietal embayment that characterizes , providing support for the phylogenetic hypothesis that and are closely related. This stepwise change of morphologic characters observed in chasmosaurine taxa that do not overlap stratigraphically is supportive of evolution by anagenesis. Recently published hypotheses that place and into separate clades are not supported. This phylogenetic relationship demonstrates unrestricted movement of large-bodied taxa between hitherto purported northern and southern provinces in the late Campanian, weakening support for the hypothesis of extreme faunal provincialism in the Late Cretaceous Western Interior.

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http://www.ncbi.nlm.nih.gov/pmc/articles/PMC7278894PMC
http://dx.doi.org/10.7717/peerj.9251DOI Listing

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