Systematic revision of the . complex: Resurrection of the genus and introduction of the new genus .

The genus is resurrected on the basis of comparative morphology and phylogeny of partial nuc 18S rDNA (18S) and translation elongation factor 1-alpha () nucleotide sequences. The genus is characterized by the firm upper surface of the pseudoaethalium, accreted but easily separable sporothecae, a tubular or fibrous columella, and spores with a reticulate ornamentation consisting of 7-9 meshes across the diameter. In addition to the currently known single species (= ), two new members of are described: from coniferous forests of Europe, east Asia, and southeast Asia, and from temperate and subarctic regions of North America and alpine regions of Europe. A second genus, , is described to accommodate . The fruiting body of this species is transitional between a pseudoaethalium and a true aethalium. It is covered by a contiguous membranous cortex formed by the fused tips of the sporothecae, a feature typical for aethalia. However, the inner portions of sporothecae remain discernible, a feature more typical for pseudoaethalia. Columellae of are formed by perforated plates, and the spores have a reticulate ornamentation consisting of 2-5 meshes across the diameter. For , we could confirm records only for tropical regions and Japan, whereas all studied European specimens, including those mentioned in current monographs, represent species of .