Following publication of the original article [1], a typographical error in the formula for calculating d in the "Scans for local adaptation" subsection in the Method section, was identified. The correct formula should be.

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http://www.ncbi.nlm.nih.gov/pmc/articles/PMC6784328PMC
http://dx.doi.org/10.1186/s13059-019-1821-1DOI Listing

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