Revision of the Oriental species of Latreille (). Part 2: the species group (Coleoptera, Carabidae, Lebiini).

The species group of the genus Latreille from Asia (in the sense of Casale and Shi 2018) is revised with six species recognized. Four new species are described: Shi & Casale, (type locality: Taiwan: Siling, 24.65°N, 121.42°E); Casale & Shi, (type locality: Vietnam: "Cochinchina"); Shi & Casale, (type locality: Yunnan: Caiyanghe, 22.60°N, 101.12°E); and Casale & Shi, (type locality: Philippines: Nagtipunan, 16.22°N, 121.60°E). Jedlička, 1963 is confirmed as an available and valid species name, and C. Kirschenhofer, 1986 is newly synonymized with it. A phylogenetic analysis of Oriental species, based on adult morphological characters, is performed. The results show that the monophyly of most species groups in Oriental is accepted, but the group appears polyphyletic and comprises three lineages. However, because many species relationships in the cladogram lack significant supporting, presently the group was remained to use for morphological convenience. Five types of female reproductive tracts were recognized, corresponding to five branches in the cladogram.