The trace fossil Chondrites, a highly branched burrow system of unknown endobenthic deposit feeders, occurs in all types of sediment, including those deposited under anaerobic conditions. In some cases, such as the Jurassic Posidonienschiefer Formation of Germany, Chondrites occurs in black, laminated, carbonaceous sediment that was deposited in chemically reducing conditions. In other cases, such as numerous oxic clastic and carbonate units throughout the geologic column, Chondrites typically represents the last trace fossil in a biotutbation sequence. This indicates that the burrow system was produced deep within the sediment in the anaerobic zone below the surficial oxidized zone that was characterized by freely circulating and oxidizing pore waters.
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