The present concepts of evolution and species delineation in prokaryotes are considered. Recently a considerable extension of knowledge on the processes of microevolution of medically significant bacteria was noted alongside with the importance of horizontal and lateral transfer of genes. The phylophenetic concept of species was considered in detail. The inclusion of the ecological criterion into a phylophenetic concept of a species is supposed to facilitate the development of more adequate notion on the evolution of bacteria, the improvement of species delineation in prokaryotes, their classification and nomenclature.
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