Placenta and fetal membranes of the four-toed elephant shrew, Petrodromus tetradactylus.

The four-toed elephant shrew is the only extant member of the genus Petrodromus. Some features of its placentation are found in other elephant shrews and may be important in consolidating the elephant shrews in the order Macroscelidea. These include implantation in an embryo chamber at the caudal end of the uterine horn; a large yolk sac; a large, lobulated allantoic sac; a distinct layer of proliferative trophoblastic cells between the labyrinthine and spongy zones of the placenta; an accessory epitheliochorial placenta; and the formation of a mesoplacenta by the decidua basalis.