Although the monophyly of has been strongly demonstrated, the species relationships among approximately 20 species of have been difficult to determine because of the uniformity of their floral characteristics and extreme variation of their vegetative characters, often accompanied by high polyploid and aneuploid series and diverse habitats. In this study, we assembled 15 complete chloroplast genomes of species from East Asia and generated a plastome-based backbone phylogeny of the subgenus . As a proxy for nuclear phylogeny, we reconstructed the nuclear ribosomal DNA internal transcribed spacer (nrDNA ITS) phylogeny independently.
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