In slow motion: radula motion pattern and forces exerted to the substrate in the land snail (Mollusca, Gastropoda) during feeding.

The radula is the anatomical structure used for feeding in most species of Mollusca. Previous studies have revealed that radulae can be adapted to the food or the substrate the food lies on, but the real, forces exerted by this organ on substrates and the stresses that are transmitted by the teeth are unknown. Here, we relate physical properties of the radular teeth of (Müller.