Integrated phylogenomics and fossil data illuminate the evolution of beetles.

Beetles constitute the most biodiverse animal order with over 380 000 described species and possibly several million more yet unnamed. Recent phylogenomic studies have arrived at considerably incongruent topologies and widely varying estimates of divergence dates for major beetle clades. Here, we use a dataset of 68 single-copy nuclear protein-coding (NPC) genes sampling 129 out of the 193 recognized extant families as well as the first comprehensive set of fully justified fossil calibrations to recover a refined timescale of beetle evolution.

The Australian Lymexylidae (Coleoptera: Tenerionoidea) with one new genus and two genera new to Australia.

The generic classification of Lymexylidae is reexamined, particularly with reference to the Australian fauna. The transfer of the Madagascar genus and species Alcestoma serropalpoides Fairmaire (1895) from Melandryidae to Lymexylidae, proposed by Nikitsky in Nikitsky et al. (1998), is confirmed and the genus is considered to be a senior synonym of Protomelittomma Wheeler (1986), syn.

Eucinetus linskyi, a new replacement name for preoccupied Eucinetus bicolor Lawrence, 2019, non Reitter, 1887 (Insecta: Coleoptera: Eucinetidae).

The species name Eucinetus bicolor Lawrence was proposed by Lawrence (2019, p. 155) for a new coleopteran species from eastern Victoria, Australia. However, this name was already used for the Palaearctic Eucinetus bicolor Reitter (1887, p.

The extant species of the genus Omma Newman and description of Beutelius gen. nov. (Coleoptera: Archostemata: Ommatidae: Ommatinae).

The genus Omma Newman (type species Omma stanleyi Newman) is revised and redefined based on extant species, with examination of additional fossil species. As a result, only the species O. stanleyi Newman is retained in Omma and a new genus, Beutelius gen.

Australian Nitidulinae: general review with descriptions of new genera and species (Coleoptera: Nitidulidae).

The classification of the subfamily Nitidulinae is reviewed, as it applies to those species occurring within Australia and its territories. Comments are made concerning the currently recognised tribes and generic "complexes" within this subfamily, Australian genera are briefly discussed, with reference to recent and proposed changes in their classification, and several new genera and species are described. The following new taxa are described: Aenigmoronia echinodes, gen.

New species of Eucinetus and Noteucinetus from Australia (Coleoptera: Scirtoidea: Eucinetidae).

Two new species of Noteucinetus Bullians Leschen and 12 new species of Eucinetus Germar are described from continental Australia and Tasmania. A key is given to the two genera and all described Australian species. The following new species are described: Eucinetus bicolor, sp.

Basal polyphagan beetles in mid-Cretaceous amber from Myanmar: biogeographic implications and long-term morphological stasis.

The origin and early evolutionary history of polyphagan beetles have been largely based on evidence from the derived and diverse 'core Polyphaga', whereas little is known about the species-poor basal polyphagan lineages, which include Scirtoidea (Clambidae, Decliniidae, Eucinetidae, and Scirtidae) and Derodontidae. Here, we report two new species Acalyptomerus thayerae sp. nov.

The Australian Ciidae (Coleoptera: Tenebrionoidea): supplement.

An addendum to the revision of the Australian ciid fauna contains descriptions of 17 new species, information on recently described Australian species, addition of one described species new to the fauna, and corrections to the previous revision. Keys are provided to the Australian species of Cis, Notapterocis, Orthocis and Paratrichapus. The following new species are described: Cis apodemus, sp.

Lepicerus larva still unknown: a correction (Coleoptera: Lepiceridae, Phalacridae).

The larva of Lepicerus inaequalis Motschulsky was described by Lawrence et al. (2013) based on several early instars and one late instar collected among wet leaves and debris near Gamboa, Panama. The identification was based on collection of an adult Lepicerus in a similar habitat nearby and a combination of characters found in other myxophagan immatures but not known in other beetle larvae.

Review of the Australian Cyllodini (Coleoptera: Nitidulidae: Nitidulinae), with descriptions of new taxa, and notes on the genus Macleayania (Nitidulini).

The genera of the nitiduline tribe Cyllodini occurring in Australia are reviewed and a key is provided for their separation. Each genus is discussed, with an emphasis on the Australian fauna, and an hypothesis is given concerning the source of the introduced Camptodes species and its connection with the program for biological control of Opuntia cacti. The specimens identified by Kirejtshuk (2003) as Macleayania amphotiformis (Reitter, 1880b) were found to represent two widely overlapping species later named by Olliff (1886) in the trogossitid genus Ancyrona.

Whirling in the late Permian: ancestral Gyrinidae show early radiation of beetles before Permian-Triassic mass extinction.

Background: Gyrinidae are a charismatic group of highly specialized beetles, adapted for a unique lifestyle of swimming on the water surface. They prey on drowning insects and other small arthropods caught in the surface film. Studies based on morphological and molecular data suggest that gyrinids were the first branch splitting off in Adephaga, the second largest suborder of beetles.

The Australian Ciidae (Coleoptera: Tenebrionoidea): A Preliminary Revision.

The Australian ciid fauna is revised based on 75 described species (56 of them new) and three undescribed species, placed in 22 genera (nine of them new and nine newly recorded from Australia). The revision is considered preliminary, since about 50 other undescribed Australian species have been seen (usually as uniques or in short series), and it is likely that more will be found with increased intensive collecting. One additional New Caledonian species is also described, and one Cis species is considered to be based on a mislabelled Hawaiian specimen.

Trachelostenini sensu novo: redescriptions of Trachelostenus Solier, Myrmecodema Gebien and Leaus Matthews & Lawrence, based on adults and larvae, and descriptions of three new species of Leaus (Coleoptera: Tenebrionidae).

The former family Trachelostenidae is returned to tribal status in Tenebrionidae-Tenebrioninae and reconstituted to include Trachelostenus Solier, 1851 of southern Chile, currently in a separate family Trachelostenidae, Myrmecodema Gebien, 1943 of central Chile, currently in Stenochiinae-Cnodalonini, and Leaus Matthews & Lawrence, 1992 of eastern Australia, currently in Tenebrioninae-Titaenini. The three genera are redescribed in both the adult and larval stages. Three new species: Leaus tropicalis sp.

Review of the genera of Mycetophagidae (Coleoptera: Tenebrionoidea) with descriptions of new genera and a world generic key.

The world genera of Mycetophagidae are reviewed and four new genera described. Several adult features of Mycetophagidae are discussed, and some newly discovered secondary sexual characters are described and illustrated. A key is given to the adults of all mycetophagid genera.

Antillipeltis, a new genus of Antillean Trogossitidae (Coleoptera: Cleroidea) with a key to the Cleroidea.

Antillipeltis gen. nov. is described based on the following six new extant species and two new fossil species from Hispaniola and Puerto Rico: A.

Lectotype designations and nomenclatural changes in Xylographus Mellié (Coleoptera, Ciidae).

We designate lectotypes and propose nomenclatural changes in Xylographus Mellié (Coleoptera, Ciidae) based on type specimens deposited in the Museum of Comparative Zoology (USA), Museum für Naturkunde Berlin (Germany), the Natural History Museum (UK), Muséum d'Histoire Naturelle de la Ville de Genève (Switzerland), Muséum National d'Histoire Naturelle (France), Naturhistoriska Riksmuseet (Sweden) and Naturhistorisches Museum Wien (Austria). We designate lectotypes for the following species: Cis fultoni Broun, 1886, Xylographus anthracinus Mellié, 1849, X. bicolor Pic, 1916, X.

A Possible Larva of Lepicerus inaequalis Motschulsky (Coleoptera: Myxophaga: Lepiceridae) from Panama.

A larva is described which is presumed to be that of Lepicerus inaequalis Motschulsky based on several probable first instars and one later instar collected a few miles from and in a similar habitat to adult specimens of this species. The association is additionally based on several features also occurring in other known myxophagan larvae. A key is provided comparing these larvae with those of the other three families of Myxophaga.

Lycomimus, a new genus of Australian Ptilodactylidae (Coleoptera: Byrrhoidea).

A new genus of Ptilodactylidae, Lycomimus gen. n., based on L.

Globorentonium, a new genus of rentoniine Trogossitidae (Coleoptera: Cleroidea) from Australia and Brazil.

A new genus of Rentoniini, Globorentonium gen. n., is described including three new species: G globulum sp.

The Australian Byrrhinae (Coleoptera: Byrrhidae) with descriptions of new genera and species.

The generic classification of the Australian Byrrhinae is revised and the following new genera are described: Notolioon gen. n., Nothochaetes gen.

Loranthophila, a new genus of Australian Lyctinae (Coleoptera: Bostrichidae) associated with Mistletoe.

Loranthophila gen. n. is described, based on Minthea acanthacollis (Carter & Zeck), and comparisons are made between this genus and other members of the bostrichid subfamily Lyctinae.

Synopsis of Falsocis Pic (Coleoptera, Ciidae), new species, new records and an identification key.

Three new species of Falsocis Pic are described: Falsocis aquiloniussp. n. from Panamá, Costa Rica and Colombia, Falsocis egregiussp.