A CSSL (chromosome segment substitution line), SG-64, carrying a segment of chromosome 4 from African cultivated rice (CG-14) in the genetic background of var. Wuyujing-7 (japonica), showed a spreading panicle, which was different significantly from that of Wuyujing-7 with an erect compact panicle. The gene controlling a spreading panicle is referred to as Spr3, and is mapped on chromosome 4.
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