Adaptive evolution of flight in butterflies.

The diversity of flying animals suggests that countless combinations of flight morphologies and behaviors have evolved with specific lifestyles, thereby exploiting diverse aerodynamic mechanisms. How morphology, flight behavior, and aerodynamic properties together diversify with contrasting ecology remains to be elucidated. We studied the adaptive codivergence in wing shape, flight behavior, and aerodynamic efficiency among butterflies living in different forest strata by combining high-speed videography in the field with morphometric analyses and aerodynamic modeling.