Discovery of the fossil true bug species Urochela (Urochela) cf. melaina Zhang, 1989 (Hemiptera: Heteroptera: Urostylididae) from Japan, suggesting a wide distribution of the U. (U.) quadrinotata (Reuter, 1881) species group in East Asia in the past.

The Early Miocene lacustrine deposit (24.1 ± 2.4 Ma) of Sado Island, Niigata Prefecture, has preserved the fossil fauna of the time when the Palaearctic realm of Japan was a part of the Eurasian Continent.