Publications by authors named "Helena Shaverdo"

The group is the second species group of the New Guinean representatives of the recently described genus Shaverdo et al., 2023. The group is mainly defined by distinct scale- and/or spinula-like surface structures of the dorsal sclerite of the median lobe.

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Herein, (type species: W.J. Macleay, 1871) is described for a distinctive lineage of predominantly Australasian species previously assigned to Erichson, 1832.

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Detailed information about the known species groups of Broun, 1886 from New Guinea is presented, including species numbers, distribution, and references of species-group diagnoses, keys to the species, and species descriptions. An identification key to all species groups is provided. Phylogeny and morphological character evolution are discussed.

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The larvae of 12 species of Copelatinae, Copelatus alternatus Sharp, 1882, C. caelatipennis princeps Young, 1963, C. glyphicus (Say, 1823), C.

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Seven new species of the genus Broun, 1886 are described from three different mountain ranges of New Guinea: , , , and from the Foja Mountains; from the Cyclops Mountains; and from Wano Land. All of them are placed into the group based on the structure of their male genitalia. The species are characteristic dytiscid elements of the fauna of northern cost and the western part of central orogen of New Guinea.

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Background: The New Guinean archipelago has been shaped by millions of years of plate tectonic activity combined with long-term fluctuations in climate and sea level. These processes combined with New Guinea's location at the tectonic junction between the Australian and Pacific plates are inherently linked to the evolution of its rich endemic biota. With the advent of molecular phylogenetics and an increasing amount of geological data, the field of New Guinean biogeography begins to be reinvigorated.

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The first account of the genus Erichson, 1832 in the Solomon Islands is provided, reporting 10 species for the Archipelago. Six of these are new to science: , , , and from Guadalcanal and , and from Bougainville. Guignot, 1942, described from Tulaghi Island of the Solomons, is recorded from Guadalcanal and Santa Isabel for the first time.

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Three new species of the genus Broun, 1886: , and are described from New Guinea. The former two species are placed into the group, while the latter is suggested to be a member of a separate lineage, the newly introduced group. The only other species of that group is Shaverdo & Balke, 2014.

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Two new species of the genus Broun, 1886: and are described from New Guinea and placed into the group based on the structure of their male genitalia. The two species are very similar with respect to their external morphology and characterised by almost identical, strongly modified male antennae. However, they can easily be separated by the shape and setation of the median lobe and paramere.

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The genus in Australia is revised and nine species are recognised. One new species, , is described from Papua New Guinea (Central Province) and Cape York Peninsula (Iron Range NP and Mt Tozer). Watts, 1978, , is considered a junior synonym of Guignot, 1956, described from New Guinea.

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Nine new species groups of Broun, 1886 from New Guinea are introduced with keys to their representatives. Four groups are monotypic and include three new species: the group, the group: , the group: , and the group: The remaining five species groups include 18 species with 12 new species and one new subspecies: the group: , , , and ; the group: , , and ; the group: ; the group: ; and the group: , , , and Diagnoses of five already described species of these groups are provided, as well as comparatives notes on all species. (Balke, 1998) is a junior synonym of (Balke, 1998).

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Nineteen new species of Broun, 1886 from New Guinea are described herein: , , , , , , , , , , , , , , , , , , and . All of them, together with five already described species, have been united into the newly defined -group, a polyphyletic complex of related species with similar shape of the median lobe and paramere setation. An identification key to all known species of the group is provided, and important diagnostic characters (habitus, color, male protarsomeres 4-5, median lobes, and parameres) are illustrated.

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The habitat template concept applied to a freshwater system indicates that lotic species, or those which occupy permanent habitats along stream courses, are less dispersive than lentic species, or those that occur in more ephemeral aquatic habitats. Thus, populations of lotic species will be more structured than those of lentic species. Stream courses include both flowing water and small, stagnant microhabitats that can provide refuge when streams are low.

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Two new species and one new subspecies of Broun, 1886 from New Guinea are described: Shaverdo & Balke, , Shaverdo & Balke, , and Shaverdo & Balke, These and two already described species are assigned to the -group, which is morphologically (based on setation of the paramere) and phylogenetically close to the -group. On the latter, morphological and taxonomic notes are provided. An identification key to all known species of the groups is presented, and important diagnostic characters are illustrated.

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Six new species of New Guinea Broun, 1886 are described in this paper: , , , , , and . Although different morphologically, together with (Balke, 1998), (Balke & Hendrich, 2001), and Shaverdo, Panjaitan & Balke, 2016, they are found to form a monophyletic clade and be closely related to representatives of the -group, based on preliminary analysis of sequence data. An identification key to the species is provided, and important diagnostic characters are illustrated.

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Ten new species of Broun, 1886 from New Guinea are described: , , , , , , , , , and . All of them together with five already described species are united into the newly defined -group (with -subgroup), a polyphyletic complex of related species with lateral setation on the median lobe. In the light of newly available material, all previously described species of the -group are considered to belong to a single species, (Balke, 1998), which is now placed into the -group, and three new synonyms are therefore proposed: (Balke, 2001) .

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Exocelina manokwariensis sp. n. from West Papua is placed into the Exocelina ekari-group based on the structure of its male genitalia.

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The fate of newly settled dispersers on freshly colonized oceanic islands is a central theme of island biogeography. The emergence of increasingly sophisticated methods of macroevolutionary pattern inference paves the way for a deeper understanding of the mechanisms governing these diversification patterns on lineages following their colonization of oceanic islands. Here we infer a comprehensive molecular phylogeny for Melanesian Exocelina diving beetles.

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Early studies on Melanesian mountain systems provided insights for fundamental evolutionary and ecological concepts. These island-like systems are thought to provide opportunities in the form of newly formed, competition-free niches. Here we show that a hyperdiverse radiation of freshwater arthropods originated in the emerging central New Guinea orogen, out of Australia, about 10 million years ago.

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Exocelina baliem sp. n. is described from the Baliem Valley in the Central Mountain Range of New Guinea (Papua Province, Indonesia).

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The Exocelina ekari-group is here introduced and defined mainly on the basis of a discontinuous outline of the median lobe of the aedeagus. The group is known only from New Guinea (Indonesia and Papua New Guinea). It contained four species to date: Exocelina astrophallus (Balke, 1998), Exocelina atowaso (Shaverdo, Sagata & Balke, 2005), Exocelina munaso (Shaverdo, Sagata & Balke, 2005), and Exocelina polita (Sharp, 1882).

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