Metabolic Adjustment of High Intertidal Alga to the Tidal Cycle Includes Oscillations of Soluble Carbohydrates, Phlorotannins, and Citric Acid Content.

The brown alga is one of the species successfully adapted to intertidal conditions. Inhabiting the high intertidal zone, spends most of its life exposed to air, where it is subjected to desiccation, light, and temperature stresses. However, the physiological and biochemical mechanisms allowing this alga to tolerate such extreme conditions are still largely unknown.