Functionally referential alarm calls have stimulus specificity, distinct acoustic structure, and elicit different escape responses that are appropriate to the threat. The mechanisms by which escape responses are evoked are not fully understood and may range from eliciting innate responses to conveying representational information. White-tailed ptarmigan (Lagopus leucurus) are a long-lived alpine tundra grouse, which are preyed upon by aerial and terrestrial predators.
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