biovar () is an untypical pathogen causing a fatal anthrax-like disease in a variety of wildlife species in African rainforest areas. In contrast to and most species of the group, all strains of the cluster contain a 22 kb insertion in the gene which encodes the essential late sporulation sigma factor σ. This insertion is excised during sporulation in a site-specific recombination process resulting in an intact gene and a circular molecule.
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