N and N'-mediated recognition confers resistance to tomato brown rugose fruit virus.

Tomato brown rugose fruit virus (ToBRFV) is an emerging tobamovirus that overcomes the resistance gene used in commercial tomato plants to protect against tobamoviruses. In this article, we show that ToBRFV is recognised through its P50 replicase fragment by the resistance gene in , which triggers a hypersensitive response (HR). We also demonstrate that the gene provides protection against ToBRFV through recognition of the viral coat protein without triggering a typical HR in .