Fish fins do not contain muscles, yet fish can change their shape with high precision and speed to produce large and complex hydrodynamic forces-a combination of high morphing efficiency and high flexural stiffness that is rare in modern morphing and robotic materials. These "flexo-morphing" capabilities are rare in modern morphing and robotic materials. The thin rays that stiffen the fins and transmit actuation include mineral segments, a prominent feature whose mechanics and function are not fully understood.
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